| 1990 | - | Biological signals as handicaps - Grafen | :: | 20 |
| 1975 | - | Mate selection---a selection for a handicap - Zahavi | :: | 17 |
| 1995 | - | Communication in discrete action-response games - Hurd | :: | 10 |
| 1930 | - | The Genetical Theory of Natural Selection - Fisher | :: | 9 |
| 1998 | - | A continuous evolutionary simulation model of the attainability of honest signalling equilibria - Bullock | :: | 5 |
| 1982 | - | Heritable true fitness and bright birds: A role for parasites - Hamilton,Zuk | :: | 3 |
| 1994 | - | Sexual Selection - Andersson | :: | 3 |
| 1991 | - | The evolution of costly mate preferences II - Iwasa,Pomiankowski,Nee | :: | 2 |
| 1976 | - | Sexual selection and the handicap principle - Maynard-Smith | :: | 2 |
| 1996 | - | Why the peacock's tail is so short: Limits to sexual selection - Werner | :: | 1 |
| 1980 | - | Mate choice increases a component of offspring fitness in fruit flies - Partridge | :: | 1 |
| 1986 | - | The handicap mechanism of sexual selection does not function - Kirkpatrick | :: | 1 |
| 1973 | - | Territoriality and non-random mating in sage grouse - Wiley | :: | 1 |
| 1978 | - | The Evolution of Sex - Maynard-Smith | :: | 1 |