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Grafen, A. and Johnstone, R. A. (1993) Why we need ESS signalling theory. Philosophical Transactions: Biological Sciences, 340(1292):245--250.

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2003-Common language or Tower of Babel? On the evolutionary dynamics of signals and their meanings - van Baalen,Jansen ::
Grafen, A. & Johnstone, R. A. 1993 Why we need ESS signalling theory. Phil. Trans. R. Soc. Lond. B 340, 245--251.